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| Description: Whatever its cause, stream organisms drift downstream in large numbers. Why doesn't drift eventually eliminate organisms from the upstream sections of streams? Karl Muller (1954, 1974) hypothesized that drift would eventually wash entire populations out of streams unless organisms actively moved upstream to compensate for drift. He proposed that stream populations are maintained through a dynamic interplay between downstream and upstream dispersal that he called the colonization cycle. The colonization cycle is a dynamic view of stream populations in which upstream and downstream dispersal, as well as reproduction, have major influences on stream populations
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